In the now-popular tourist destination of Radiator Springs, the rusty old tow truck Mater has a number of roles, including proprietor of Tow Mater Towing and Salvage, grand storyteller and the heart and soul of the town.
But Mater's favorite title is best friend to Lightning McQueen. Mater values each and every dent he's earned on his escapades with Lightning over the years, and anxiously awaits further fun with his friend when Lightning returns with his latest Piston Cup.
Cheerful, honest and loyal to a fault, Mater rushes to his best friend's defense, which ultimately lands the off-duty Lightning in an international racing exhibition: the World Grand Prix. When Lightning invites Mater to come along as a member of his pit crew, Mater leaves Carburetor County for the first time ever.
Not long after their arrival in Tokyo, Mater learns that the world outside of Radiator Springs is decidedly different and its newfangled ways may take some getting used to. But when he's mistaken for an American secret agent, he falls into an intriguing adventure of his own. Caught between supporting his best friend on a globe-trotting racing competition and his new role in international espionage, Mater ends up in an explosive chase through the streets of Tokyo and Europe alongside a team of British spies and international bad guys.